Manicules, or, those pointing hands in manuscripts

Our Bodman Collection copy of Filipo Beroaldo’s Orationes: Prelectio[n]es, Praefationes: & Quaedam Mythic[ae] Historae Philippi Beroaldi (Parrhisiis, in aedibus Ascensianis: & Joannis Parui [1515]), call number HON SPCL BODMAN PA 8475 .B6 1515, is full of whimsical manicules and cartoons drawn in the margins by at least one 16th century reader. For students of Renaissance books, these marginal notations are charming; each one is drawn with a unique sleeve style. Was this reader a student of fashion as well?
Two sample pages:
closer view: